Classification of Arachnida (and other animals)Back to Arachnologia

Notes on Systematics and Taxonomy

I'll be putting stuff in here about:-Back to Opiliones Notes Of course, there are other better web sites and pages devoted more specifically to aspects of biological classification. For example there's good information on Taxonomy: Classifying Life in Kimball's Biology Pages.

Some basics of biological classification

Just for now, what is that wee thing in this picture here?..

That, basically, is the question asked by a biologist applying systematics to classify an organism. It may be a specimen taken during an ecological sampling program and the information could contribute to an assessment of biodiversity (clearly important in conservation terms) or it may be an item in a taxonomic study, concerned with providing information of direct application in the classification of more individual organisms.
Unknown specimen
We'd probably need to take a closer look, like this perhaps:
Now we can see a bit more and it would appear to be some kind of fish. With some idea of the kind of organism under examination, we could then look in much greater detail to see particular characters which would allow us to identify the species to which it belongs. Quite possibly, we could use a dichotomous key in which the presence/absence or precise form of characters to make decisions and work through a series of such decisions and eventually decide to which species he specimen belongs. The whole range of possible features can provide taxonomic information. The most obvious sort of character, of course, is structural or anatomical - "What does it look like?" - but all kinds of features could be used, even aspects of behaviour, habitat preference, geographical distribution, and so on. Physiology and biochemistry can provide useful information and with modern technology the direct sequencing of DNA can help.
This is the practical application of taxonomy - the science of biological classification, generating names for coherent groups of organisms and arranging these groups in relation to each other.
dendrogramBiologists use hierarchical classification, which is best appreciated as a dendrogram (from the Greek 'dendros' meaning branch) - a branching diagram showing the relationships:
As you can see here, group A can be divided into groups B and C, and group B also comprises groups D and E. This is looking at the dendrogram in a divisive fashion (splitting up the groups); alternatively, you could work upwards through the dendrogram, lumping groups together (agglomerative approach). Thus, in Th. latter approach we can say that D and E clump together into group C, which together with group B forms group A.

Of course, a biological classification makes most sense when we have some idea of the evolutionary relationships between the groups - this is called phylogeny, the evolutionary development of groups of organisms.
A modern approach is cladistics, describing the descendency of one group from another (that is a clade). Taxonomists look for special characters which provide evidence for such close links (these characters are called synapomorphic - from the Greek 'synapo-' = joining). cladogramThis approach generates a cladogram, in which recency of common descent is the criterion for linking groups. Care is needed, for a character to be a true synapomorph it must have common ancestry and be essentially equivalent in the two groups (another term for this is 'homologous'). False conclusions can be drawn if a character is symplesiomorphic, i.e. is just a character which has been "left over" from an earlier ancenstral group, or if it is the result of convergent evolution, i.e. looks similar but has separate origins (= 'analogous') in which case the two groups don't actually share the character but just have two different characters which look similar. A plesiomorph is a primitive character, in contrast to a apomorph which is a derived (or advanced, modified) character. A neat outline of cladistics has been written by  Lynne M. Clos, or ther's a good set of definitions for cladistics by M. Tevfik Dorak (with lots of good links to other relevant pages).

For a good application of cladistics to spiders see Chapter 10 of Foelix Rainer's book Biology of Spiders, 2nd. ed., published by Oxford University Press in 1996.

fancy cladogram/phylogenyAn artistically-drawn cladistics diagram may look something like this,
more like a phylogeny, which is not too different from the old (mid-19th century) family trees representing classificatory schemes.

(Not really too different either in general form from our little fish up above, which in its overall shape could be taken for a phylogenetic diagram - handy as a wee logo for this page!)

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